腋芽萌发能力对再生稻产量影响的研究进展

doi:10.16819/j.1001-7216.2020.9097

摘要/Abstract

摘要：

再生稻能够充分利用温光资源,一种两收,能提高复种指数和单位面积粮食产量,是增加水稻总产的一条重要途径,对保障我国粮食安全意义重大。再生季有效穗数是影响再生稻产量的重要因子,而有效穗数取决于再生芽萌发能力。提高再生稻腋芽萌发能力对提高再生季水稻产量有关键作用。本文介绍了国内再生稻发展现状,综述了腋芽萌发能力和其影响因素以及提高腋芽能力的技术措施等,并提出了未来加强再生稻腋芽萌发的研究重点,以期为再生稻高产栽培提供理论依据。

关键词: 再生稻, 腋芽, 萌发能力, 产量

Abstract:

Ratoon rice can make full use of heat and light resources. It has the advantage of two harvests with one sowing, and it can increase multiple cropping index and grain yield per unit area, which has great importance to rice production and food security in our country. The effective panicle number plays a vital role in the yield formation in ratoon rice, which depends on the germination ability of regeneration buds. Therefore, improving the axillary bud germination ability of ratoon rice holds the key to increasing rice yield in regenerative season. In this paper, we outlined the current research advances in the mechanism of axillary bud germination and the influencing factors of the germination ability as well as the technical measures to improve axillary bud germination ability. We also proposed the research highlights in the future on enhancing axillary bud germination. It will lay a theoretical basis for higher yield in ratoon rice.

Key words: ratoon rice, axillary bud, germination ability, yield

中图分类号:

S511.01

汪浩, 张强, 张文地, 李思宇, 黄健, 朱安, 刘立军. 腋芽萌发能力对再生稻产量影响的研究进展[J]. 中国水稻科学, 2020, 34(3): 205-216.

Hao WANG, Qiang ZHANG, Wendi ZHANG, Siyu LI, Jian HUANG, An ZHU, Lijun LIU. Advances in the Effects of the Ability of Axillary Bud Germination on Grain Yield in Ratoon Rice[J]. Chinese Journal OF Rice Science, 2020, 34(3): 205-216.

图/表 3

参考文献 82

[1]	Dong H, Chen Q, Wang W, Peng S, Huang J, Cui K, Nie L X.The growth and yield of a wet-seeded rice-ratoon rice system in central China[J]. Field Crops Research, 2017, 208: 55-59.
[2]	Qian C, He A, Wang W, Peng S, Huang J, Cui K.Comparisons of regeneration rate and yields performance between inbred and hybrid rice cultivars in a direct seeding rice-ratoon rice system in central China[J]. Field Crops Research, 2018, 223: 164-170.
[3]	林文雄, 陈鸿飞, 张志兴, 徐倩华, 屠乃美, 方长旬, 任万军. 再生稻产量形成的生理生态特性与关键栽培技术的研究与展望[J]. 中国生态农业学报,2015, 23(4): 392-401.
	Lin W X, Chen H F, Zhang Z X, Xu Q H, Tu N M, Fang C X, Ren W J.Research and prospect on physio-ecological properties of ratoon rice yield formation and its key cultivation technology[J]. Chinese Journal of Eco- Agriculture, 2015, 23(4): 392-401. (in Chinese with English abstract)
[4]	何花榕, 翁国华, 郭灵灵, 张丹. 影响再生稻腋芽萌发因素的研究进展[J]. 福建稻麦科技, 2008(3): 43, 61-63.
	He H R, Weng G H, Guo L L, Zhang D. Research progress on factors affecting the germination of axillary buds in ratoon rice[J]. Fujian Science & Technology of Rice & Wheat, 2008(3): 43, 61-63. (in Chinese with English abstract)
[5]	胡香玉, 钟旭华, 彭碧琳, 刘彦卓, 黄农荣, 梁开明, 潘俊峰, 傅友强. 不同氮肥运筹下低桩机收再生稻的产量和经济效益[J]. 中国稻米, 2019(4): 1-7.
	Hu X Y, Zhong X H, Peng B L, Liu Y Z, Huang N R, Liang K M, Pan J F, Fu Y Q.Yield and economic benefits of low-pile harvesting of reclaimed rice under different nitrogen fertilizer operations[J]. China Rice, 2019(4): 1-7. (in Chinese with English abstract)
[6]	Lin W X.Developmental status and problems of rice ratooning[J]. Journal of Integrative Agriculture, 2019, 18(1): 246-247.
[7]	陈基旺, 帅泽宇, 屠乃美, 易镇邪. 湖南再生稻发展现状与对策分析[J]. 中国稻米, 2018, 24(5): 68-72.
	Chen J W, Shuai Z Y, Tu N M, Yi Z X.Analysis on the status and countermeasures of Hunan regeneration rice development[J]. China Rice, 2018, 24(5): 68-72. (in Chinese with English abstract)
[8]	段秀建, 张巫军, 姚雄, 李经勇, 唐永群, 佘小明, 肖人鹏. 杂交中稻机收蓄留再生稻高产高效栽培技术[J]. 杂交水稻, 2019, 34(1): 44-46.
	Duan X J, Zhang W J, Yao X, Li J Y, Tang Y Q, She X M, Xiao R P.High-yielding and high-efficiency cultivation techniques for ratooning rice of mid-season hybrid rice under mechanical harvest conditions[J]. Hybrid Rice, 2019, 34(1): 44-46. (in Chinese with English abstract)
[9]	吴延寿, 姚晓云, 曹国军, 陈春莲, 熊运华, 尹建华. 优质再生稻产量形成和稻米品质比较分析[J]. 杂交水稻, 2019, 34(5): 57-63.
	Wu Y S, Yao X Y, Cao G J, Chen C L, Xiong Y H, Yin J H.Comparative analysis of yield formation and rice quality of high quality ratoon rice[J]. Hybrid Rice, 2019, 34(5): 57-63. (in Chinese with English abstract)
[10]	徐富贤, 熊洪, 张林, 朱永川, 蒋鹏, 郭晓艺, 刘茂. 再生稻产量形成特点与关键调控技术研究进展[J]. 中国农业科学, 2015, 48(9): 1702-1717.
	Xu F X, Xiong H, Zhang L, Zhu Y C, Jiang P, Guo X Y, Liu M.Research on the characteristics of yield formation and key regulation techniques of ratoon rice[J]. Scientia Agricultura Sinica, 2015, 48(9): 1702-1717. (in Chinese with English abstract)
[11]	蒲莉芳, 向晓霞. 不同播种期对C两优34156再生稻产量及其构成因素的影响[J]. 杂交水稻, 2018, 33(5): 39-41.
	Pu L F, Xiang X X.Effects of different sowing dates on yield and its components of C Liangyou 34156 ratoon rice[J]. Hybrid Rice, 2018, 33(5): 39-41. (in Chinese with English abstract)
[12]	晏月明, 王绪信, 邵启明, 刘保国, 任昌福. 水稻亚种间杂交再生力特性的遗传[J]. 西南农业大学学报, 1992(6): 517-521.
	Yan Y M, Wang X X, Shao Q M, Liu B G, Ren C F.Heredity of reproductive characteristics of intersubspecific hybrids in rice[J]. Journal of Southwest Agricultural University, 1992(6): 517-521. (in Chinese with English abstract)
[13]	任天举, 李经勇, 张晓春, 王培华, 余官平, 卢瑶. 水稻杂种F1再生力特性的配合力和遗传力分析[J]. 西南农业学报, 2004(3): 287-291.
	Ren T J, Li J Y, Zhang X C, Wang P H, Yu G P, Lu Y.Analysis of combining ability and heritability of regenerative characteristics of F1 hybrids in rice[J]. Southwest China Journal of Agricultural Sciences, 2004(3): 287-291. (in Chinese with English abstract)
[14]	蒋彭炎. 水稻高产栽培理论与技术讲座(14): 再生稻的生育特性与高产栽培要点[J]. 中国稻米, 1996(6): 30-33.
	Jiang P Y.The Theory and techniques of high-yield cultivation of rice (14): The growth characteristics and high-yield cultivation points of ratooning rice[J]. China Rice, 1996(6): 30-33. (in Chinese with English abstract)
[15]	谭震波, 沈利爽, 陆朝福, 陈英, 朱立煌, 周开达, 袁祚廉. 水稻再生能力和头季稻产量性状的QTL定位及其遗传效应分析. 作物学报, 1997(3): 289-295.
	Tan Z B, Shen L S, Lu C F, Chen Y, Zhu L H, Zhou K D, Yuan Z L.QTL mapping and genetic effect analysis of rice germination ability and yield traits of main crop rice. Acta Agronomica Sinica, 1997(3): 289-295. (in Chinese with English abstract)
[16]	郑景生, 李义珍, 林文雄. 应用SSR标记定位水稻再生力和再生产量及其构成的QTL. 分子植物育种, 2004(3): 342-347.
	Zheng J S, Li Y Z, Lin W X.Identification of QTL for ratooning ability and grain yield traits in ratoon rice based on SSR marker.Molecular Plant Breeding, 2004 (3): 342-347. (in Chinese with English abstract)
[17]	杨川航, 王玉平, 涂斌, 李婷, 胡亮, 李仕贵. 利用RIL群体对水稻再生力及相关农艺性状的QTL分析. 作物学报, 2012, 38(7): 1240-1246.
	Yang C H, Wang Y P, Tu B, Li T, Hu L, Li S G.QTL Analysis of rice ratooning ability and related agronomic traits using RIL population.Acta Agronomica Sinica, 2012, 38(7): 1240-1246. (in Chinese with English abstract)
[18]	李兴星, 郑剑, 周军杰, 杨永清, 秦小健, 张汉马, 梁永书. 利用籼粳交“热粳35/协B”F2群体对水稻再生力的QTL分析.分子植物育种, 2016, 14(9): 2383-2391.
	Li X X, Zheng J, Zhou J J, Yang Y Q, Qin X J, Zhang H M, Liang Y S.QTL analysis of rice regenerative ability using the Rejing 35/Xie B F2 population of indica-japonica cross.Molecular Plant Breeding, 2016, 14(9): 2383-2391. (in Chinese with English abstract)
[19]	李阳. 不同品种中稻蓄留再生稻产量形成的差异与再生芽化学调控初探[D]. 武汉: 华中农业大学, 2014.
	Li Y.Differences in yield formation and regenerative bud chemical regulation of rice grown in different rice varieties[D]. Wuhan: Huazhong Agricultural University, 2014. (in Chinese with English abstract)
[20]	习敏, 吴文革, 汪靖桂, 汪宏伟, 陈刚, 许有尊. 再生稻种植产量差形成的研究[J]. 华北农学报, 2017, 32(1): 104-110.
	Xi M, Wu W G, Wang J G, Wang H W, Chen G, Xu Y Z.Studies on the formation of yield differences in ratooning rice planting[J]. Acta Agriculturae Boreali-sinica, 2017, 32(1): 104-110. (in Chinese with English abstract)
[21]	李贵勇, 谭丽明, 刘玉文, 黄洁, 陈双云, 夏琼梅, 龙瑞平, 杨从党. 再生稻的产量形成及其高产群体分析[J]. 江西农业大学学报, 2015, 37(4): 577-583.
	Li G Y, Tan L M, Liu Y W, Huang J, Chen S Y, Xia Q M, Long R P, Yang C D.Analysis of the yield formation and high-yield population of ratooning rice[J]. Acta Agriculturae Universitatis Jiangxiensis, 2015, 37(4): 577-583. (in Chinese with English abstract)
[22]	吴芸紫, 段门俊, 刘章勇, 陈阜, 金涛. 播种期对3个再生稻品种产量及产量构成因子的影响[J]. 作物杂志, 2017(2): 151-156.
	Wu Y Z, Duan M J, Liu Z Y, Chen F, Jin T.Effects of sowing date on yield and yield components of three ratoon rice varieties[J]. Crops, 2017(2): 151-156. (in Chinese with English abstract)
[23]	鲁远源, 任天举. 川东地区中稻-再生稻生产及其影响因素的关联分析[J]. 西南农业学报, 1994(3): 49-53.
	Lu Y Y, Ren T J.Correlation analysis of medium rice-regenerate rice production and its influencing factors in Eastern Sichuan[J]. Southwest China Journal of Agricultural Sciences, 1994 (3): 49-53. (in Chinese with English abstract)
[24]	郑常敏. 再生稻生长发育及产量构成特点初探[J]. 中国稻米, 1996(4): 11-12.
	Zheng C M.Preliminary study on characteristics of growth and development and yield components of ratooning rice[J]. China Rice, 1996(4): 11-12. (in Chinese with English abstract)
[25]	丁卫东, 罗瑾. 隆回县再生稻品种筛选及分[J]. 作物研究, 2009, 23(2): 89-91.
	Ding W D, Luo J.Screening and classification of ratoon rice varieties in Longhui County[J]. Crop Research, 2009, 23(2): 89-91. (in Chinese with English abstract)
[26]	罗丽华, 肖应辉, 刘国华, 唐文帮, 陈立云. 不同类型水稻再生特性的比较研究[J]. 广西农业科学, 2006(2): 123-126.
	Luo L H, Xiao Y H, Liu G H, Tang W B, Chen L Y.Comparative research on regeneration characteristics of different types of rice[J]. Guangxi Agricultural Science, 2006(2): 123-126. (in Chinese with English abstract)
[27]	周文新. 不同类型再生稻生育特性及源库关系比较研究[D]. 长沙: 湖南农业大学, 2006.
	Zhou W X.Comparative study on the growth characteristics and source-sink relationship of different types of ratoon rice[D]. Changsha: Hunan Agricultural University, 2006. (in Chinese with English abstract)
[28]	段门俊, 吴芸紫, 田玉聪, 刘永武, 刘章勇, 陈阜, 金涛. 不同品种再生稻产量及品质比较研究[J]. 作物杂志, 2018(2): 61-67.
	Duan M J, Wu Y Z, Tian Y C, Liu Y W, Liu Z Y, Chen F, Jin T.Comparative study on yield and quality of different varieties of ratooning rice[J]. Crops, 2018(2): 61-67. (in Chinese with English abstract)
[29]	李华平. 杂交稻甬优1540作再生稻种植表现与高产栽培技术[J]. 福建农业科技, 2019(8): 54-56.
	Li H P.Performance and high-yield cultivation techniques of hybrid rice yongyou 1540 as ratooning rice[J]. Fujian Agricultural Science and Technology, 2019 (8): 54-56. (in Chinese with English abstract)
[30]	熊渠, 刘丹, 王记安, 程建平, 李拥军, 王兰. 甬优4949再生稻全程机械化种植表现及高产栽培技术[J]. 湖北农业科学, 2018, 57(9): 16-19.
	Xiong Q, Liu D, Wang J A, Cheng J P, Li Y J, Wang L.Performance and high-yield cultivation technology of the whole mechanized planting of Yongyou 4949 ratoon rice[J]. Hubei Agricultural Sciences, 2018, 57(9): 16-19. (in Chinese with English abstract)
[31]	廖海林, 郑景生, 李小萍, 解振兴, 黄信恩, 李义珍, 黄育民, 王侯聪. 2个再生稻品种机械化生产的产量构成分析[J]. 厦门大学学报: 自然科学版, 2016, 55(6): 853-859.
	Liao H L, Zheng J S, Li X P, Xie Z X, Huang X E, Li Y Z, Huang Y M, Wang H C.Analysis on the yield composition of mechanized production of two ratoon rice varieties[J]. Journal of Xiamen University: Natural Science, 2016, 55(6): 853-859. (in Chinese with English abstract)
[32]	林强, 姜照伟, 林祁, 王颖姮, 张初长, 卓传营, 谢鸿光, 蒋家焕, 谢华安, 张建福. 杂交稻强再生力鉴定指标的建立及组合筛选[J]. 福建农业学报, 2019, 34(8): 873-882.
	Lin Q, Jiang Z W, Lin Q, Wang Y H, Zhang C C, Zhuo C Y, Xie H G, Jiang J H, Xie H A, Zhang J F.Establishment and combination screening of identification indexes for strong regeneration of hybrid rice[J]. Fujian Journal of Agricultural Sciences, 2019, 34(8): 873-882. (in Chinese with English abstract)
[33]	任天举, 张晓春, 王培华, 李经勇. 杂交中稻、再生稻两季高产组合的主要特征特性及配合力效应[J]. 西南农业学报, 2005(4): 382-386.
	Ren T J, Zhang X C, Wang P H, Li J Y.The Main characteristics and combining ability effects of two-season high-yield combinations of hybrid mid-season and ratooning rice[J]. Southwest China Journal of Agricultural Sciences, 2005(4): 382-386. (in Chinese with English abstract)
[34]	王直华. 杂交中稻半期旱作对再生稻生长发育的影响[D]. 武汉: 华中农业大学, 2009.
	Wang Z H.The effects of half-period dryland cultivation on the growth and development of ratooning rice[D]. Wuhan: Huazhong Agricultural University, 2009. (in Chinese with English abstract)
[35]	陈其敏. 水氮密调控对黄华占再生稻产量及氮肥利用率的影响[D]. 长沙: 湖南农业大学, 2017.
	Chen Q M.Effect of water and nitrogen density regulation on Huanghuazhan's ratoon rice yield and nitrogen fertilizer utilization rate[D]. Changsha: Hunan Agricultural University, 2017. (in Chinese with English abstract)
[36]	任天举, 蒋志成, 王培华, 李经勇, 张晓春, 鲁远源, 刘贤双. 杂交中稻再生力与头季稻农艺性状的相关性研究[J]. 作物学报, 2006(4): 613-617.
	Ren T J, Jiang Z C, Wang P H, Li J Y, Zhang X C, Lu Y Y, Liu X S.Correlation of ratooning ability with its main crop agronomic traits in mid-season hybrid rice[J]. Acta Agronomica Sinica, 2006(4): 613-617. (in Chinese with English abstract)
[37]	刘爱中, 张胜文, 屠乃美. 稻桩贮藏同化产物的分配与再生稻腋芽再生率及产量构成的关系[J]. 华北农学报, 2008(3): 190-193.
	Liu A Z, Zhang S W, Tu N M.The relationship between the distribution of assimilation products in rice storage and the regeneration rate and yield components of regenerated rice aphid[J]. Acta Agriculturae Boreali- Sinica, 2008(3):190-193. (in Chinese with English abstract)
[38]	梁成山, 杨玉仙, 侯松德. 杂交稻再生稻生物学特性及高产栽培技术措施探索[J]. 农业科技通讯, 2018(11): 216-219.
	Liang C S, Yang Y X, Hou S D.Exploration of biological characteristics and high-yield cultivation techniques of hybrid rice in hybrid rice[J]. Agricultural Science and Technology Communication, 2018(11): 216-219. (in Chinese with English abstract)
[39]	陈鸿飞, 庞晓敏, 张仁, 张志兴, 徐倩华, 方长旬, 李经勇, 林文雄. 不同水肥运筹对再生季稻根际土壤酶活性及微生物功能多样性的影响[J]. 作物学报, 2017, 43(10): 1507-1517.
	Chen H F, Pang X M, Zhang R, Zhang Z X, Xu Q H, Fang C X, Li J Y, Lin W X.Effects of different water and fertilizer applications on soil enzyme activities and microbial functional diversity in regenerated rice rhizosphere[J]. Acta Agronomica Sinica, 2017, 43(10): 1507-1517. (in Chinese with English abstract)
[40]	徐富贤, 熊洪, 朱永川, 张林, 郭晓艺, 刘茂. 促芽肥施用时期对不同源库类型杂交中稻再生力的影响[J]. 杂交水稻, 2010, 25(3): 57-63, 99.
	Xu F X, Xiong H, Zhu Y C, Zhang L, Guo X Y, Liu M.Effects of germination-promoting period on regenerative capacity of hybrid rice in different source-sink types[J]. Hybrid Rice, 2010, 25(3): 57-63, 99. (in Chinese with English abstract)
[41]	陈鸿飞, 张志兴, 林文雄. 促芽肥对水稻再生芽萌发生长过程蛋白质表达的影响[J]. 中国生态农业学报, 2014, 22(12): 1405-1413.
	Chen H F, Zhang Z X, Lin W X.Effects of nitrogen application for bud development on protein expression of ratooning buds of rice[J]. Chinese Journal of Eco-Agriculture, 2014, 22(12): 1405-1413. (in Chinese with English abstract)
[42]	胡志华, 李大明, 徐小林, 黄庆海, 柳开楼, 胡惠文, 叶会财, 周利军, 余喜初. 再生稻轻简化种植技术研究进展[J]. 中国稻米, 2017, 23(3): 13-17.
	Hu Z H, Li D M, Xu X L, Huang Q H, Liu K L, Hu H W, Ye H C, Zhou L J, Yu X C.Research progress on light and simplified planting techniques of reclaimed rice[J]. China Rice, 2017, 23(3): 13-17. (in Chinese with English abstract)
[43]	蒋廷杰, 易镇邪, 屠乃美. 留桩高度对培矮64S/E32再生特性的影响[J]. 湖南农业大学学报: 自然科学版, 2005(4): 359-363.
	Jiang T J, Yi Z X, Tu N M. Effects of pile height on the regeneration characteristics of Pei'ai 64S/E32[J]. Journal of Hunan Agricultural University: Natural Science, 2005(4): 359-363. (in Chinese with English abstract)
[44]	刘爱中, 邹冬生, 屠乃美, 刘飞. 留桩高度对再生稻生长发育及产量的影响[J]. 安徽农业科学, 2007(17): 5120-5121, 5172.
	Liu A Z, Zou D S, Tu N M, Liu F.Effects of pile height on growth and yield of ratooning rice[J]. Anhui Agricultural Sciences, 2007(17): 5120-5121, 5172. (in Chinese with English abstract)
[45]	练红, 周海涛, 陈维建, 李广华, 李木进, 周光怀, 盛建华, 李红霞, 肖灿霞, 徐一兰. 播种期和留桩高度对再生稻产量及产量构成因素的影响[J]. 湖南农业科学, 2017(4): 28-31.
	Lian H, Zhou H T, Chen W J, Li G H, Li M J, Zhou G H, Sheng J H, Li H X, Xiao C X, Xu Y L.Effects of sowing date and height of piles on yield and yield components of ratoon rice[J]. Hunan Agricultural Sciences, 2017(4): 28-31. (in Chinese with English abstract)
[46]	易镇邪, 周文新, 屠乃美. 留桩高度对再生稻源库性状与物质运转的影响[J]. 中国水稻科学, 2009, 23(5): 509-516.
	Yi Z X, Zhou W X, Tu N M.Effects of stubble height of the main crop on source-sink characteristics and assimilates transportation in ratooning rice[J]. Chinese Journal of Rice Science, 2009, 23(5): 509-516. (in Chinese with English abstract)
[47]	熊洪, 方文. 再生稻腋芽萌发与产量形成的生态研究[J]. 生态学报, 1994(2): 161-167.
	Xiong H, Fang W.Ecological study on germination and yield formation of alfalfa buds in ratoon rice[J]. Acta Ecologica Sinica, 1994(2): 161-167. (in Chinese with English abstract)
[48]	黄新杰. 再生稻根芽萌发对母体营养与激素的响应[D]. 长沙: 湖南农业大学, 2012.
	Huang X J.Response of root bud germination of ratoon rice to maternal nutrition and hormones[D]. Changsha: Hunan Agricultural University, 2012. (in Chinese with English abstract)
[49]	江世华, 熊洪, 方文, 罗文质. 四川省再生稻高产综合栽培技术研究[J]. 西南农业大学学报, 1995(3): 189-192.
	Jiang S H, Xiong H, Fang W, Luo W Z.Research on the High-yield integrated cultivation techniques of ratoon rice in Sichuan Province[J]. Journal of Southwest Agricultural University, 1995 (3): 189-192. (in Chinese with English abstract)
[50]	肖应辉. 水稻再生特性及其与内源激素等生理因素的关系[D]. 长沙: 湖南农业大学, 2002.
	Xiao Y H.Relationship between rice regeneration characteristics and physiological factors such as endogenous hormones[D]. Changsha: Hunan Agricultural University, 2002. (in Chinese with English abstract)
[51]	贾信荣. 不同时期喷施赤霉素对再生稻芽苗及产量的影响[J]. 现代农业科技, 2015(14):124-125.
	Jia X R.Effects of spraying gibberellin on regeneration rice sprouts and yield in different periods[J]. Modern Agricultural Science and Technology, 2015(14): 124-125. (in Chinese with English abstract)
[52]	杨文钰, 马均, 杨世民, 袁继超, 倪甦. 杂交中稻培育再生稻施用赤霉素两季增产的机理和技术[J]. 杂交水稻, 1994(6): 18-20.
	Yang W Y, Ma J, Yang S M, Yuan J C, Ni S. The mechanism and technique of GA3 spraying to increase grain yield of both main and ratoon crops in medium hybrid rice[J]. Hybrid Rice, 1994(6): 18-20. (in Chinese with English abstract)
[53]	王光明, 刘保国, 陈静, 任昌福. 内源激素ABA对水稻再生芽萌发的影响[J]. 西南农业大学学报, 1997(4): 24-28.
	Wang G M, Liu B G, Chen J, Ren C F.Effects of endogenous hormone ABA on germination of rice buds[J]. Journal of Southwest Agricultural University, 1997(4): 24-28. (in Chinese with English abstract)
[54]	李经勇, 唐永群. 杂交水稻再生芽萌发生长与植株内源细胞分裂素含量变化的关系[J]. 杂交水稻, 2002(3): 53-55.
	Li J Y, Tang Y Q.Relationship between the length of regenerated shoot germination and the change of endogenous cytokinin content in hybrid rice[J]. Hybrid Rice, 2002(3): 53-55. (in Chinese with English abstract)
[55]	黄新杰, 屠乃美, 易镇邪, 周文新. 不同激素处理对再生稻产量及相关生理特性的影响[J]. 杂交水稻, 2017, 32(4): 71-75, 80.
	Huang X J, Tu N M, Yi Z X, Zhou W X.Effects of different hormone treatments on yield and related physiological characteristics of ratoon rice[J]. Hybrid Rice, 2017, 32(4): 71-75, 80. (in Chinese with English abstract)
[56]	徐富贤, 熊洪, 赵甘霖, 洪松. 杂交中稻强再生力品种的冠层特征研究[J]. 作物学报, 2002(3): 426-430.
	Xu F X, Xiong H, Zhao G L, Hong S.Canopy characteristics of strong regenerative varieties of hybrid mid-season rice[J]. Acta Agronomica Sinica, 2002(3): 426-430. (in Chinese with English abstract)
[57]	徐富贤, 熊洪. 杂交中稻粒叶比与再生力的关系[J]. 中国水稻科学, 2000(4): 58-61.
	Xu F X, Xiong H.Relationship between grain-leaf ratio and regenerative capacity of hybrid rice[J]. Chinese Journal of Rice Science, 2000(4): 58-61. (in Chinese with English abstract)
[58]	方立魁, 丛云飞, 刘丽, 何光华, 曾卓华. 重庆市再生稻产区主导品种筛选[J]. 贵州农业科学, 2016, 44(12): 6-9.
	Fang L K, Cong Y F, Liu L, He G H, Zeng Z H.Screening of dominant varieties of regenerated rice production areas in Chongqing[J]. Guizhou Agricultural Sciences, 2016, 44(12): 6-9. (in Chinese with English abstract)
[59]	郑小雄. 优质稻佳辐占作再生稻高产栽培技术[J]. 福建农业, 2013(2): 22-23.
	Zheng X X.High-yield cultivation techniques for high-quality rice Jiafuzhan accounted for regeneration rice[J]. Fujian Agriculture, 2013 (2): 22-23. (in Chinese with English abstract)
[60]	许晖, 徐正猛, 吴芸紫, 汪德志, 史方春, 马军, 杨洪勇, 谢磊. 节水抗旱稻旱优73在江汉平原作再生稻种植的表现及其机械化高产栽培技术[J]. 杂交水稻, 2019, 34(6): 39-40, 100.
	Xu H, Xu Z M, Wu Y Z, Wang D Z, Shi F C, Ma J, Yang H Y, Xie L.Performance and mechanized high-yielding cultivation techniques of water-saving and drought- resistant hybrid rice combination Hanyou 73 grown as ratoon rice in Jianghan Plain[J]. Hybrid Rice, 2019, 34(6): 39-40, 100. (in Chinese with English abstract)
[61]	吴集斌, 李玉白, 王大来, 赵忠桂. 再生稻不同播种期对产量的影响及其适宜的播种期[J]. 湖南环境生物职业技术学院学报, 2011, 17(1): 1-4.
	Wu J B, Li Y B, Wang D L, Zhao Z G.Effects of different sowing dates on yield of ratoon rice and its suitable sowing date[J]. Journal of Hunan Environmental Biotechnology College, 2011, 17(1): 1-4. (in Chinese with English abstract)
[62]	柳开楼, 秦江涛, 张斌. 播种期对轻简栽培方式再生稻源库关系的影响[J]. 土壤, 2012, 44(4): 686-695.
	Liu K L, Qin J T, Zhang B.Effects of sowing date on the relationship between source and sink of regenerated rice in light cultivation mode[J]. Soil, 2012, 44(4): 686-695. (in Chinese with English abstract)
[63]	Daliri M S, Eftekhari A, Mobasser H R.Effect of cutting time and cutting height on yield and yield components of ratoon rice (Tarom Langrodi Variety)[J]. Asian Journal of Plant Sciences, 2009, 8(1): 89-91.
[64]	张上守, 卓传营, 郑荣和, 李义珍. 再生稻超高产优化集成技术[J]. 中国稻米, 2007(3): 44-48.
	Zhang S S, Zhuo C Y, Zheng R H, Li Y Z.Optimized integration technology for super high yield of reclaimed rice[J]. China Rice, 2007(3): 44-48. (in Chinese with English abstract)
[65]	Hhillens Lambers D.Varietal improvement for ratooning rice: traits, procedures, collaboration//Rice Ratooning[M]. Los Banos, Philippines: International Rice Research Institute, 1987: 247-255.
[66]	Vergara B S, Lopez F S S, Chauhan J S. Morpholpgy and physiology of ratooning rice//Rice Ratooning[M]. Los Banos, Philippines: International Rice Research Institute, 1987: 31-40.
[67]	何花榕, 房贤涛, 谢祖钦, 张居念, 卓传营, 蔡光璟, 杨惠杰. 不同氮钾施用量对再生稻产量及其构成因素的影响[J]. 福建农业学报, 2016, 31(7): 699-703.
	He H R, Fang X T, Xie Z Q, Zhang J N, Zhuo C Y, Cai G J, Yang H J.Effects of different nitrogen and potassium application rates on yield and its components of ratoon rice[J]. Fujian Journal of Agricultural Sciences, 2016, 31(7): 699-703. (in Chinese with English abstract)
[68]	刘保国, 王光明, 陈静, 陈国惠, 任昌福. 促芽肥影响水稻再生芽萌发的生理生化基础[J]. 西南农业学报, 1998(S3): 45-49.
	Liu B G, Wang G M, Chen J, Chen G H, Ren C F.Physiological and biochemical basis of germination promoting germination of rice buds[J]. Southwest China Journal of Agricultural Sciences, 1998(S3): 45-49. (in Chinese with English abstract)
[69]	习敏, 徐秀娟, 吴文革, 汪靖桂, 陈再高, 孙雪原, 许有尊. 促芽肥对再生稻准两优608产量和主要品质性状的影响[J]. 中国稻米, 2018, 24(3): 93-96.
	Xi M, Xu X J, Wu W G, Wang J G, Chen Z G, Sun X Y, Xu Y Z.Effects of germination on the yield and main quality traits of regenerated rice Zhunliangyou 608[J]. China Rice, 2018, 24(3): 93-96. (in Chinese with English abstract)
[70]	冉茂林, 熊洪, 徐富贤, 张志勇. 杂交水稻生育后期光合产物的分配对再生稻的影响[J]. 绵阳经济技术高等专科学校学报, 1999(4): 6-10.
	Ran M L, Xiong H, Xu F X, Zhang Z Y.Effect of the distribution of photosynthetic products on the regeneration rice in the late growth stage of hybrid rice[J]. Journal of Mianyang University of Economics and Technology, 1999(4): 6-10. (in Chinese with English abstract)
[71]	徐富贤, 熊洪, 朱永川, 张林, 郭晓艺. 利用杂交中稻齐穗期剑叶叶绿素计读数(SPAD值)预测再生稻促芽肥高效施用量[J]. 中国水稻科学, 2009, 23(1): 51-56.
	Xu F X, Xiong H, Zhu Y C, Zhang L, Guo X Y.Estimation of efficient rate of nitrogen application for promoting ratooning bud development using chlorophyll meter reading(SPAD value) of flag leaf at the full heading stage of main crop in mid-season hybrid rice[J]. Chinese Journal of Rice Science, 2009, 23(1): 51-56. (in Chinese with English abstract)
[72]	胡香玉, 钟旭华, 梁开明, 黄农荣, 潘俊峰, 刘彦卓, 傅友强, 彭碧琳. 广东再生稻研究进展与展望[J]. 中国稻米, 2019(6): 1-5.
	Hu X Y, Zhong X H, Liang K M, Huang N R, Pan J F, Liu Y Z, Fu Y Q, Peng B L.Research progress and prospect of Guangdong regenerative rice[J]. China Rice, 2019(6): 1-5. (in Chinese with English abstract)
[73]	张荟, 郑轶, 涂诗航, 周鹏, 卓传营, 张上守. 杂交稻新品种高留桩再生力筛选试验[J]. 福建稻麦科技, 2015, 33(3): 11-14.
	Zhang H, Zheng Y, Tu S H, Zhou P, Zhuo C Y, Zhang S S.Screening test of reproductive ability of new hybrid rice varieties with high retention piles[J]. Fujian Science ＆ Technology of Rice ＆ Wheat, 2015, 33(3): 11-14. (in Chinese with English abstract)
[74]	熊洪, 冉茂林, 徐富贤, 洪松. 南方稻区再生稻研究进展及发展[J]. 作物学报, 2000(3): 297-304.
	Xiong H, Ran M L, Xu F X, Hong S.Research progress and development of ratooning rice in southern rice region[J]. Acta Agronomica Sinica, 2000 (3): 297-304. (in Chinese with English abstract)
[75]	王鸿林. 机收低留桩再生稻高产栽培技术[J]. 福建稻麦科技, 2017, 35(2): 11-13.
	Wang H L.High-yield cultivation techniques of reclaimed rice with low yield and remaining piles[J]. Fujian Science ＆ Technology of Rice ＆ Wheat, 2017, 35(2): 11-13. (in Chinese with English abstract)
[76]	邓文财. 优质稻“佳辐占”作再生稻机收低留桩高产栽培技术[J]. 福建农业科技, 2011(1): 7-8.
	Deng W C.High-quality rice "Jiafuzhan" as a regenerative rice harvester with low yield and high yield cultivation technology[J]. Fujian Agricultural Science and Technology, 2011(1): 7-8. (in Chinese with English abstract)
[77]	Wai A H, An G.Axillary meristem initiation and bud growth in rice[J]. Journal of Plant Biology, 2017, 60(5): 440-451.
[78]	徐云姬, 许阳东, 李银银, 钱希旸, 王志琴, 杨建昌. 干湿交替灌溉对水稻花后同化物转运和籽粒灌浆的影响[J]. 作物学报, 2018, 44(4): 554-56.
	Xu Y J, Xu Y D, Li Y Y, Qian X Y, Wang Z Q, Yang J C.Effects of dry-wet alternate irrigation on post-flowering assimilate transport and grain filling of rice[J]. Acta Agronomica Sinica, 2018, 44(4):554-56. (in Chinese with English abstract)
[79]	徐国伟, 吕强, 陆大克, 王贺正, 陈明灿. 干湿交替灌溉耦合施氮对水稻根系性状及籽粒库活性的影响[J]. 作物学报, 2016, 42(10): 1495-1505.
	Xu G W, Lv Q, Lu D K, Wang H Z, Chen M C.Effects of wetting and drying alter native irrigation coupling with nitrogen application on root characteristic and grain-sink activity[J]. Acta Agronomica Sinica, 2016, 42(10): 1495-1505. (in Chinese with English abstract)
[80]	陈婷婷, 许更文, 钱希旸, 王志琴, 张耗, 杨建昌. 花后轻干-湿交替灌溉提高水稻籽粒淀粉合成相关基因的表达[J]. 中国农业科学, 2015, 48(7): 1288-1299.
	Chen T T, Xu G W, Qian X Y, Wang Z Q, Zhang H, Yang J C.Post-anthesis alternative wetting and moderate soil drying irrigation enhance gene expression of enzymes involved in starch synthesis in rice grains[J]. Scientia Agricultura Sinica, 2015, 48(7):1288-1299. (in Chinese with English abstract)
[81]	陈婷婷, 褚光, 华小龙, 潘燕, 王志琴, 杨建昌. 花后干湿交替灌溉对水稻强、弱势粒蛋白质表达的影响[J]. 中国农业科学, 2013, 46(22): 4665-4678.
	Chen T T, Chu G, Hua X L, Pan Y, Wang Z Q, Yang J C.Effects of post-harvest drying and wetting alternate irrigation on protein expression of strong and weak granules in rice[J]. Scientia Agricultura Sinica, 2013, 46(22): 4665-4678. (in Chinese with English abstract)
[82]	张自常, 李鸿伟, 陈婷婷, 王学明, 王志琴, 杨建昌. 畦沟灌溉和干湿交替灌溉对水稻产量与品质的影响[J]. 中国农业科学, 2011, 44(24): 4988-4998.
	Zhang Z C, Li H W, Chen T T, Wang X M, Wang Z Q, Yang J C.Effect of furrow irrigation and alternate wetting and drying irrigation on grain yield and quality of rice[J]. Scientia Agricultura Sinica, 2011, 44(24): 4988-4998. (in Chinese with English abstract)

文献 Reference	年份 Year	材料 Material	QTL	染色体 Chromosome	标记区间 Marker interval	贡献率 Variation explained/%	加性效应 Additive effect
[15]	1997	窄叶青8号/京系17 Zhaiyeqing 8/Jingxi 17	Ra3a	3	RG369A–RG369B	9.8	0.38
			Ra4a	4	C975–RG119	16.2	-0.20
			Ra5a	5	C624–RG573	8.5	0.37
			Ra6a	6	RG653–RG433	16.5	0.49
			Ra6b	6	G294–G422	13.4	0.46
			Ra7a	7	RG650–C285	18.3	0.51
[16]	2004	明恢86/佳辐占 Minghui 86/Jiafuzhan	qSNP	1	RM1–RM283	13.6	15.27
			qSS	7	RM18–RM234	9.1	5.81
			qPN	7	RM18–RM234	8.0	1.31
			qGY	7	RM18–RM234	7.9	2.77
[17]	2012	糯89-1/蜀恢527 Nuo 89-1/Shuhui 527	qRA4	4	RM401–RM307	8.17	0.32
[17]	2012	糯89-1/蜀恢527 Nuo 89-1/Shuhui 527	qRA5	5	RM163–RM153	7.09	-0.39
[18]	2016	热粳35/协B Rejing 35/Xie B	qRa8	8	RM6976–RN264	23.35	0.19
[18]	2016	热粳35/协B Rejing 35/Xie B	qRa11	11	RM2191–RM6239	16.01	-0.13

文献 Reference	年份 Year	材料 Material	QTL	染色体 Chromosome	标记区间 Marker interval	贡献率 Variation explained/%	加性效应 Additive effect
[15]	1997	窄叶青8号/京系17 Zhaiyeqing 8/Jingxi 17	Ra3a	3	RG369A–RG369B	9.8	0.38
			Ra4a	4	C975–RG119	16.2	-0.20
			Ra5a	5	C624–RG573	8.5	0.37
			Ra6a	6	RG653–RG433	16.5	0.49
			Ra6b	6	G294–G422	13.4	0.46
			Ra7a	7	RG650–C285	18.3	0.51
[16]	2004	明恢86/佳辐占 Minghui 86/Jiafuzhan	qSNP	1	RM1–RM283	13.6	15.27
			qSS	7	RM18–RM234	9.1	5.81
			qPN	7	RM18–RM234	8.0	1.31
			qGY	7	RM18–RM234	7.9	2.77
[17]	2012	糯89-1/蜀恢527 Nuo 89-1/Shuhui 527	qRA4	4	RM401–RM307	8.17	0.32
[17]	2012	糯89-1/蜀恢527 Nuo 89-1/Shuhui 527	qRA5	5	RM163–RM153	7.09	-0.39
[18]	2016	热粳35/协B Rejing 35/Xie B	qRa8	8	RM6976–RN264	23.35	0.19
[18]	2016	热粳35/协B Rejing 35/Xie B	qRa11	11	RM2191–RM6239	16.01	-0.13

品种名称 Cultivar name	头季稻产量 Yield of main crop/(t·hm^-2)	再生季产量 Yield of ratoon rice/(t·hm^-2)	两季总产 Total yield/(t·hm^-2)
两优6326 Liangyou 6326	8.80	6.62	15.4
天两优616 Tianliangyou 616	8.50	6.38	14.9
广两优476 Guangliangyou 476	8.41	6.57	15.0
丰两优香1号 Fengliangyouxiang 1	8.45	6.19	14.6
平均值 Average	8.54	6.37	14.9

品种名称 Cultivar name	头季稻产量 Yield of main crop/(t·hm^-2)	再生季产量 Yield of ratoon rice/(t·hm^-2)	两季总产 Total yield/(t·hm^-2)
两优6326 Liangyou 6326	8.80	6.62	15.4
天两优616 Tianliangyou 616	8.50	6.38	14.9
广两优476 Guangliangyou 476	8.41	6.57	15.0
丰两优香1号 Fengliangyouxiang 1	8.45	6.19	14.6
平均值 Average	8.54	6.37	14.9

品种类型 Type	品种名称 Combination name	单株分蘖数 Tiller number per plant	成熟期叶面积指数 LAI at maturity	齐穗至成熟干物质积累量 DMA/(t·hm^-2)	每穗粒数 Spikelet number per panicle	再生季/头季稻产量比 YR
适宜作再生稻种植 Suitable for ratoon rice	两优6326、天两优616、广两优476、丰两优香1号 Liangyou 6236, Tianliangyou 616 Guangliangyou 476, Fengliangyouxiang 1	12.0–13.2	4.9–6.2	6.2–8.2	150.1–169.0	0.70–0.78
不适作再生稻种植 Not suitable for ratoon rice	C两优018、E两优476、两优0815、盛泰优018 C liangyou 018, E liangyou 476 Liangyou 0815, Shengtaiyou 018	10.7–11.5	4.3–4.5	4.4–5.4	114.9–136.4	0.55–0.62